About the Authors

Craig R. White

Affiliation Centre for Ornithology, School of Biosciences, The University of Birmingham, Birmingham, United Kingdom

Norman Day

Affiliation Centre for Ornithology, School of Biosciences, The University of Birmingham, Birmingham, United Kingdom

Patrick J. Butler

Affiliation Centre for Ornithology, School of Biosciences, The University of Birmingham, Birmingham, United Kingdom

Graham R. Martin

To whom correspondence should be addressed. E-mail: g.r.martin@bham.ac.uk

Affiliation Centre for Ornithology, School of Biosciences, The University of Birmingham, Birmingham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CW PB GM. Performed the experiments: CW GM ND. Analyzed the data: CW. Wrote the paper: CW PB GM ND.