About the Authors

Martin R. Miller

To whom correspondence should be addressed. E-mail: m.r.miller@sheffield.ac.uk

Affiliation Department of Animal and Plant Sciences, University of Sheffield, Sheffield, United Kingdom

Andrew White

Affiliation Department of Mathematics and the Maxwell Institute for Mathematical Sciences, Heriot-Watt University, Edinburgh, Scotland

Kenneth Wilson

Affiliation Department of Biological Sciences, Lancaster Environment Centre, Lancaster University, Lancaster, United Kingdom

Michael Boots

Affiliation Department of Animal and Plant Sciences, University of Sheffield, Sheffield, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: MM. Wrote the paper: MB MM AW KW. Other: Conceived and designed the model: MB AW MM. Interpretation of the results: KW MB AW MM. Performed the computer simulations: MM AW.