About the Authors

Heather M. Scobie

Contributed equally to this work with: Heather M. Scobie, John M. Marlett

Affiliations Infectious Disease Laboratory, The Salk Institute for Biological Studies, La Jolla, California, United States of America, Cellular and Molecular Biology Graduate Program, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

John M. Marlett

Contributed equally to this work with: Heather M. Scobie, John M. Marlett

Affiliation Infectious Disease Laboratory, The Salk Institute for Biological Studies, La Jolla, California, United States of America

G. Jonah A. Rainey

Current address: Macrogenics, Inc., Rockville, Maryland, United States of America

Affiliation Infectious Disease Laboratory, The Salk Institute for Biological Studies, La Jolla, California, United States of America

D. Borden Lacy

Affiliation Department of Microbiology and Immunology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

R. John Collier

Affiliation Department of Microbiology and Molecular Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

John A.T. Young

To whom correspondence should be addressed. E-mail: jyoung@salk.edu

Affiliations Infectious Disease Laboratory, The Salk Institute for Biological Studies, La Jolla, California, United States of America, Cellular and Molecular Biology Graduate Program, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Department of Microbiology and Immunology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Department of Microbiology and Molecular Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

JATY and RJC hold equity in PharmAthene Inc. (Baltimore, MD).

Author Contributions

Conceived and designed the experiments: GR JY HS. Performed the experiments: HS JM. Analyzed the data: RC GR JY HS JM DL. Wrote the paper: GR JY HS JM.