About the Authors

Angus M. Tester

Contributed equally to this work with: Angus M. Tester, Jennifer H. Cox

Affiliation University of British Columbia Centre for Blood Research, Departments of Oral Biological and Medical Sciences, and Biochemistry and Molecular Biology, University of British Columbia, Vancouver, British Columbia, Canada

Jennifer H. Cox

Contributed equally to this work with: Angus M. Tester, Jennifer H. Cox

Affiliation University of British Columbia Centre for Blood Research, Departments of Oral Biological and Medical Sciences, and Biochemistry and Molecular Biology, University of British Columbia, Vancouver, British Columbia, Canada

Andrea R. Connor

Affiliation University of British Columbia Centre for Blood Research, Departments of Oral Biological and Medical Sciences, and Biochemistry and Molecular Biology, University of British Columbia, Vancouver, British Columbia, Canada

Amanda E. Starr

Affiliation University of British Columbia Centre for Blood Research, Departments of Oral Biological and Medical Sciences, and Biochemistry and Molecular Biology, University of British Columbia, Vancouver, British Columbia, Canada

Richard A. Dean

Affiliation University of British Columbia Centre for Blood Research, Departments of Oral Biological and Medical Sciences, and Biochemistry and Molecular Biology, University of British Columbia, Vancouver, British Columbia, Canada

Xose S. Puente

Affiliation Department Bioquimica y Biologia Molecular, Instituto Universitario de Oncologia, Universidad de Oviedo, Oviedo, Spain

Carlos López-Otín

Affiliation Department Bioquimica y Biologia Molecular, Instituto Universitario de Oncologia, Universidad de Oviedo, Oviedo, Spain

Christopher M. Overall

To whom correspondence should be addressed. E-mail: chris.overall@ubc.ca

Affiliation University of British Columbia Centre for Blood Research, Departments of Oral Biological and Medical Sciences, and Biochemistry and Molecular Biology, University of British Columbia, Vancouver, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CL CO XP. Performed the experiments: AT JC AC AS RD. Analyzed the data: AT JC. Contributed reagents/materials/analysis tools: CL XP. Wrote the paper: CO AT JC.