About the Authors

Jonas Binladen

Contributed equally to this work with: Jonas Binladen, M. Thomas P. Gilbert

Affiliation Center for Ancient Genetics, Institute of Biology, University of Copenhagen, Copenhagen, Denmark

M. Thomas P. Gilbert

Contributed equally to this work with: Jonas Binladen, M. Thomas P. Gilbert

Affiliation Center for Ancient Genetics, Institute of Biology, University of Copenhagen, Copenhagen, Denmark

Jonathan P. Bollback

Affiliation Center for Bioinformatics and Institute of Biology, University of Copenhagen, Copenhagen, Denmark

Frank Panitz

Affiliation Department of Genetics and Biotechnology, Danish Institute of Agricultural Sciences Research Centre Foulum, Tjele, Denmark

Christian Bendixen

Affiliation Department of Genetics and Biotechnology, Danish Institute of Agricultural Sciences Research Centre Foulum, Tjele, Denmark

Rasmus Nielsen

Affiliation Center for Bioinformatics and Institute of Biology, University of Copenhagen, Copenhagen, Denmark

Eske Willerslev

To whom correspondence should be addressed. E-mail: ewillerslev@bi.ku.dk

Affiliation Center for Ancient Genetics, Institute of Biology, University of Copenhagen, Copenhagen, Denmark

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RN EW MG JB. Performed the experiments: MG JB CB FP. Analyzed the data: RN MG JB JB. Contributed reagents/materials/analysis tools: EW MG JB JB CB. Wrote the paper: EW MG JB. Other: Conceived initial idea to apply 5′ tagged primers to the GS20 during discussion on the topic with MTPG: JB.