About the Authors

Hiroshi Okabe

Affiliations Department of Pathology, School of Medicine, University of California San Francisco, San Francisco, California, United States of America, UCSF Comprehensive Cancer Center, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Sang-Hyun Lee

Affiliations Cancer Research Institute, School of Medicine, University of California San Francisco, San Francisco, California, United States of America, UCSF Comprehensive Cancer Center, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Janyaporn Phuchareon

Affiliations Department of Pathology, School of Medicine, University of California San Francisco, San Francisco, California, United States of America, UCSF Comprehensive Cancer Center, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Donna G. Albertson

Affiliations Cancer Research Institute, School of Medicine, University of California San Francisco, San Francisco, California, United States of America, UCSF Comprehensive Cancer Center, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Frank McCormick

Affiliations Cancer Research Institute, School of Medicine, University of California San Francisco, San Francisco, California, United States of America, UCSF Comprehensive Cancer Center, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Osamu Tetsu

To whom correspondence should be addressed. E-mail: tetsu@cc.ucsf.edu

Affiliations Department of Pathology, School of Medicine, University of California San Francisco, San Francisco, California, United States of America, UCSF Comprehensive Cancer Center, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: OT HO. Performed the experiments: OT HO SL JP. Analyzed the data: FM OT HO SL JP. Contributed reagents/materials/analysis tools: FM DA OT HO. Wrote the paper: OT.