About the Authors

Yoshitake Sano

Contributed equally to this work with: Yoshitake Sano, Tadashi Nakaya

Affiliation RIKEN, Brain Science Institute, Behavioral Genetics, Wako, Japan

Tadashi Nakaya

Contributed equally to this work with: Yoshitake Sano, Tadashi Nakaya

Affiliation Laboratory of Neuroscience, Graduate School of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Steve Pedrini

Affiliation Farber Institute for Neurosciences of Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Shizu Takeda

Affiliation RIKEN, Brain Science Institute, Behavioral Genetics, Wako, Japan

Kanae Iijima-Ando

Affiliations Farber Institute for Neurosciences of Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Koichi Iijima

Affiliations Farber Institute for Neurosciences of Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Paul M. Mathews

Affiliation Dementia Research Program, The Nathan S. Kline Institute for Psychiatric Research, Orangeburg, New York, United States of America

Shigeyoshi Itohara

Affiliation RIKEN, Brain Science Institute, Behavioral Genetics, Wako, Japan

Sam Gandy

To whom correspondence should be addressed. E-mail: samgandy@earthlink.net (SG); tsuzuki@pharm.hokudai.ac.jp (TS)

Affiliation Farber Institute for Neurosciences of Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Toshiharu Suzuki

To whom correspondence should be addressed. E-mail: samgandy@earthlink.net (SG); tsuzuki@pharm.hokudai.ac.jp (TS)

Affiliation Laboratory of Neuroscience, Graduate School of Pharmaceutical Sciences, Hokkaido University, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TS KI KI SI. Performed the experiments: SP PM TN KI KI YS ST. Analyzed the data: SG SP TS TN KI KI SI YS ST. Contributed reagents/materials/analysis tools: PM TS. Wrote the paper: SG TS KI KI SI.