About the Authors

Don Klinkenberg

To whom correspondence should be addressed. E-mail: d.klinkenberg@vet.uu.nl

Affiliation Theoretical Epidemiology, Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, Netherlands

Christophe Fraser

Affiliation Department of Infectious Disease Epidemiology, Imperial College London, London, United Kingdom

Hans Heesterbeek

Affiliation Theoretical Epidemiology, Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CF HH DK. Performed the experiments: DK. Analyzed the data: DK. Wrote the paper: CF HH DK