About the Authors

Kurt J. Williams

williamsk@dcpah.msu.edu

Affiliation Department of Pathobiology and Diagnostic Investigation, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

N. Edward Robinson

Affiliation Large Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Ailam Lim

Affiliation Diagnostic Center for Population and Animal Health, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Christina Brandenberger

Affiliation Department of Pathobiology and Diagnostic Investigation, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Roger Maes

Affiliation Department of Pathobiology and Diagnostic Investigation, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Ashley Behan

Affiliation Large Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Steven R. Bolin

Affiliation Department of Pathobiology and Diagnostic Investigation, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Competing Interests

This study was funded by Pfizer Animal Health, New York, N.Y. There are no other declarations to be made related to employment, consultancy, patents, products in development etc. related to this manuscript. The authors' relationship with Pfizer Animal Health does not alter their adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KJW SRB NER AB RM AL. Performed the experiments: KJW SRB. Analyzed the data: KJW SRB RM AL CB. Contributed reagents/materials/analysis tools: KJW SRB CB. Wrote the manuscript: KJW SRB. Clinical management of animals: NER AB.