About the Authors

Maria I. Zervou

Affiliation Laboratory of Molecular Medicine and Human Genetics, Medical School of Crete, Heraklion, Crete, Greece

Efsevia Myrthianou

Affiliation Laboratory of Molecular Medicine and Human Genetics, Medical School of Crete, Heraklion, Crete, Greece

Irene Flouri

Affiliation Department of Rheumatology, Clinical Immunology and Allergy, Medical School of Crete, Heraklion, Greece

Darren Plant

Affiliation NIHR Manchester Musculoskeletal Biomedical Research Unit, Manchester Academy of Health Sciences, Manchester, United Kingdom

Gregory Chlouverakis

Affiliation Department of Social Medicine, University of Crete, Heraklion, Greece

Francesc Castro-Giner

Affiliation Molecular and Population Genetics Laboratory, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Panayiota Rapsomaniki

Affiliation Department of Rheumatology, Clinical Immunology and Allergy, Medical School of Crete, Heraklion, Greece

Anne Barton

Affiliations NIHR Manchester Musculoskeletal Biomedical Research Unit, Manchester Academy of Health Sciences, Manchester, United Kingdom, Arthritis Research UK Epidemiology Unit, University of Manchester, Manchester, United Kingdom

Dimitrios T. Boumpas

Affiliations Medical School, National and Kapodistrian University of Athens, Athens, Greece, Biomedical Research Foundation of the Academy of Athens, Athens, Greece, Institute of Molecular Biology and Biotechnology, Foundation of Research and Technology–Hellas, Heraklion, Greece

Prodromos Sidiropoulos

Contributed equally to this work with: Prodromos Sidiropoulos, George N. Goulielmos

Affiliation Department of Rheumatology, Clinical Immunology and Allergy, Medical School of Crete, Heraklion, Greece

George N. Goulielmos

Contributed equally to this work with: Prodromos Sidiropoulos, George N. Goulielmos

goulielmos@med.uoc.gr

Affiliation Laboratory of Molecular Medicine and Human Genetics, Medical School of Crete, Heraklion, Crete, Greece

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GNG DP AB IF. Performed the experiments: MIZ EM IF PS PR DP. Analyzed the data: GNG MIZ DP GC PS DTB FCG. Contributed reagents/materials/analysis tools: PS DTB GNG PR AB FCG. Wrote the manuscript: GNG DP MIZ AB GC DTB FCG.