About the Authors

Mirjam Frischknecht

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Helena Niehof-Oellers

Affiliation Düsseldorf, Germany

Vidhya Jagannathan

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Marta Owczarek-Lipska

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Cord Drögemüller

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Elisabeth Dietschi

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Gaudenz Dolf

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Bernd Tellhelm

Affiliation Department of Veterinary Clinical Science, Small Animal Clinic, Justus-Liebig-University, Giessen, Germany

Johann Lang

Affiliation Department of Clinical Veterinary Medicine, Division of Clinical Radiology, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Katriina Tiira

Affiliations Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences and Department of Medical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Hannes Lohi

Affiliations Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences and Department of Medical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Tosso Leeb

tosso.leeb@vetsuisse.unibe.ch

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Bern, Switzerland

Competing Interests

H. Lohi owns shares of the company Genoscoper, which currently offers COL11A2 genotyping as a commercial genetic test. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MF TL. Performed the experiments: MF HN-O VJ MO-L CD BT. Analyzed the data: MF VJ CD GD JL TL. Contributed reagents/materials/analysis tools: HN-O ED BT KT HL. Wrote the paper: MF JL TL.