About the Authors

Sharon Elaine Warner

swarner@rvc.ac.uk

Affiliation Structure and Motion Laboratory, Comparative Biomedical Sciences, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Phillip Pickering

Affiliation Structure and Motion Laboratory, Comparative Biomedical Sciences, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Olga Panagiotopoulou

Affiliation Structure and Motion Laboratory, Comparative Biomedical Sciences, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Thilo Pfau

Affiliation Veterinary Clinical Sciences, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Lei Ren

Affiliation School of Mechanical, Aerospace and Civil Engineering, The University of Manchester, Manchester, Greater Manchester, United Kingdom

John Richard Hutchinson

Affiliation Structure and Motion Laboratory, Comparative Biomedical Sciences, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Edited the final manuscript: SEW PP OP TP LR JRH. Conceived and designed the experiments: JRH OP. Performed the experiments: SEW TP LR. Analyzed the data: SEW. Contributed reagents/materials/analysis tools: PP. Wrote the paper: SEW.