About the Authors

Danielle A. Badro

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Bouchra Douaihy

Contributed equally to this work with: Bouchra Douaihy, Marc Haber

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Marc Haber

Contributed equally to this work with: Bouchra Douaihy, Marc Haber

Affiliations The Lebanese American University, Chouran, Beirut, Lebanon, Institut de Biologia Evolutiva (CSIC-UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Sonia C. Youhanna

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Angélique Salloum

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Michella Ghassibe-Sabbagh

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Brian Johnsrud

Affiliation Modern Thought and Literature, Stanford University, Stanford, California, United States of America

Georges Khazen

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Elizabeth Matisoo-Smith

Affiliation Allan Wilson Centre for Molecular Ecology and Evolution, University of Otago, Dunedin, New Zealand

David F. Soria-Hernanz

Affiliations Institut de Biologia Evolutiva (CSIC-UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain, The Genographic Project, National Geographic Society, Washington, DC, United States of America

R. Spencer Wells

Affiliation The Genographic Project, National Geographic Society, Washington, DC, United States of America

Chris Tyler-Smith

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Daniel E. Platt

Affiliation Computational Biology Centre, IBM TJ Watson Research Centre, Yorktown Heights, New York, United States of America

Pierre A. Zalloua

pierre.zalloua@lau.edu.lb

Affiliations The Lebanese American University, Chouran, Beirut, Lebanon, Harvard School of Public Health, Boston, Massachusetts, United States of America

The Genographic Consortium

Membership of the The Genographic Consortium is provided in the Acknowledgments section.

Competing Interests

IBM has no marketed products associated with the contents of this paper. IBM has no income derived from contracts related to consultancy or capital or intellectual assets in this research. The affiliation of an author (DP) with IBM does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: PZ CTS RSW EMS. Performed the experiments: DB BD SY AS. Analyzed the data: DP MH DSH GK. Contributed reagents/materials/analysis tools: BJ MGS. Wrote the paper: PZ DP DB.