About the Authors

Helen M. Alexander

halexander@ku.edu

Affiliation Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas, United States of America

Aaron W. Reed

Affiliation School of Biological Sciences, University of Missouri-Kansas City, Kansas City, Missouri, United States of America

W. Dean Kettle

Affiliation Kansas Biological Survey, University of Kansas, Lawrence, Kansas, United States of America

Norman A. Slade

Affiliations Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas, United States of America, Natural History Museum/Biodiversity Research Center, University of Kansas, Lawrence, Kansas, United States of America

Sarah A. Bodbyl Roels

Affiliation Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas, United States of America

Cathy D. Collins

Affiliation Department of Biology, Colby College, Waterville, Maine, United States of America

Vaughn Salisbury

Affiliation Kansas Biological Survey, University of Kansas, Lawrence, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HA WDK NAS VS. Performed the experiments: HA WDK SABR CDC VS. Analyzed the data: HA AWR NAS. Contributed reagents/materials/analysis tools: WDK NAS AWR. Wrote the paper: HA AWR NAS SABR CDC.