About the Authors

Lewis G. Halsey

l.halsey@roehampton.ac.uk

Affiliation Department of Life Sciences, University of Roehampton, London, United Kingdom

David A. R. Watkins

Affiliation Department of Life Sciences, University of Roehampton, London, United Kingdom

Brendan M. Duggan

Affiliation Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

Co-author Lewis G. Halsey is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LH DW. Performed the experiments: LH DW. Analyzed the data: LH DW BD. Contributed reagents/materials/analysis tools: LH. Wrote the paper: LH DW BD.