About the Authors

Kristin N. Harper

kh2383@columbia.edu

Affiliation Department of Environmental Health Sciences, Columbia University Medical Center, New York, New York, United States of America

Robert D. Fyumagwa

Affiliation Wildlife Veterinary Program, Tanzania Wildlife Research Institute, Arusha, Tanzania

Richard Hoare

Affiliation Wildlife Veterinary Program, Tanzania Wildlife Research Institute, Arusha, Tanzania

Philemon N. Wambura

Affiliation Department of Veterinary Microbiology and Parasitology, Faculty of Veterinary Medicine, Sokoine University of Agriculture, Morogoro, Tanzania

Dorian H. Coppenhaver

Affiliation Department of Microbiology and Immunology, University of Texas Medical Branch, Galveston, Texas, United States of America

Robert M. Sapolsky

Affiliations Biological Sciences, Stanford University, Stanford, California, United States of America, Institute of Primate Research, National Museums of Kenya, Nairobi, Kenya

Susan C. Alberts

Affiliations Institute of Primate Research, National Museums of Kenya, Nairobi, Kenya, Department of Biology, Duke University, Durham, North Carolina, United States of America

Jenny Tung

Affiliations Department of Evolutionary Anthropology, Duke University, Durham, North Carolina, United States of America, Population Research Institute, Duke University, Durham, North Carolina, United States of America

Jeffrey Rogers

Affiliation Human Genome Sequencing Center, Baylor College of Medicine, Houston, Texas, United States of America

Morris Kilewo

Affiliation Department of Ecology, Tanzania National Parks, Arusha, Tanzania

Emmanuel K. Batamuzi

Affiliation Department of Veterinary Surgery and Theriogenology, Sokoine University of Agriculture, Morogoro, Tanzania

Fabian H. Leendertz

Affiliation Research Group Emerging Zoonoses, Robert Koch-Institute, Berlin, Germany

George J. Armelagos

Affiliation Department of Anthropology, Emory University, Atlanta, Georgia, United States of America

Sascha Knauf

Affiliation Pathology Unit, German Primate Centre, Leibniz Institute for Primate Research, Göttingen, Germany

Competing Interests

This project received support from the Scil Animal Care Company, Telinject Inc., and Translogistic Inc. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KNH RDF RH SK GJA. Analyzed the data: KNH FHL SK GJA. Contributed reagents/materials/analysis tools: KNH FHL SK GJA. Wrote the paper: KNH RDF SK. Involved in collecting the samples: RDF RH PNW DHC RMS SCA JT JR MK EKB FHL SK. Carried out the serological and genetic analyses: KNH FHL SK GJA.