About the Authors

Alexa A. Pragman

Affiliation Department of Medicine, University of Minnesota, Minneapolis, Minnesota, United States of America

Hyeun Bum Kim

Current address: Department of Biomedical Sciences, Cummings School of Veterinary Medicine, Tufts University, North Grafton, Massachusetts, United States of America

Affiliation Department of Veterinary and Biomedical Sciences, University of Minnesota, St. Paul, Minnesota, United States of America

Cavan S. Reilly

Affiliation Division of Biostatistics, School of Public Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Christine Wendt

Affiliation Department of Medicine, VA Medical Center, Minneapolis, Minnesota, United States of America, and for the FORTE Study Group

Richard E. Isaacson

isaac015@umn.edu

Affiliation Department of Veterinary and Biomedical Sciences, University of Minnesota, St. Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AAP CW REI. Performed the experiments: AAP. Analyzed the data: AAP REI CW CR. Contributed reagents/materials/analysis tools: HBK CW. Wrote the paper: AAP HBK CW REI.