About the Authors

Wan Yang

Affiliations Department of Civil and Environmental Engineering, Virginia Tech, Blacksburg, Virginia, United States of America, Department of Environmental Health Sciences, Mailman School of Public Health, Columbia University, New York, New York, United States of America

Subbiah Elankumaran

Affiliation Department of Biomedical Sciences and Pathobiology, Virginia-Maryland Regional College of Veterinary Medicine, Virginia Tech, Blacksburg, Virginia, United States of America

Linsey C. Marr

lmarr@vt.edu

Affiliation Department of Civil and Environmental Engineering, Virginia Tech, Blacksburg, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WY LCM. Performed the experiments: WY. Analyzed the data: WY LCM SE. Contributed reagents/materials/analysis tools: SE. Wrote the paper: WY LCM.