About the Authors

Ben Wielstra

ben.wielstra@naturalis.nl

Affiliations Naturalis Biodiversity Center, Leiden, The Netherlands, University of Twente, Faculty of Geo-Information Science and Earth Observation – ITC, Enschede, The Netherlands

Wouter Beukema

Affiliation University of Twente, Faculty of Geo-Information Science and Earth Observation – ITC, Enschede, The Netherlands

Jan W. Arntzen

Affiliation Naturalis Biodiversity Center, Leiden, The Netherlands

Andrew K. Skidmore

Affiliation University of Twente, Faculty of Geo-Information Science and Earth Observation – ITC, Enschede, The Netherlands

Albertus G. Toxopeus

Affiliation University of Twente, Faculty of Geo-Information Science and Earth Observation – ITC, Enschede, The Netherlands

Niels Raes

Affiliation Naturalis Biodiversity Center, Leiden, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BW WB JWA AKS AGT NR. Performed the experiments: BW WB. Analyzed the data: BW WB. Wrote the paper: BW.