About the Authors

Vivian L. de Oliveira

Affiliation Laboratory of Medical Immunology, Department of Laboratory Medicine, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Romy R. M. C. Keijsers

Affiliations Laboratory of Medical Immunology, Department of Laboratory Medicine, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands, Dermatology Department, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Peter C. M. van de Kerkhof

Affiliation Dermatology Department, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Marieke M. B. Seyger

Affiliation Dermatology Department, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Esther Fasse

Affiliation Laboratory of Medical Immunology, Department of Laboratory Medicine, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Lars Svensson

Affiliation Department of Disease Pharmacology, LEO Pharma, Ballerup, Denmark

Markus Latta

Affiliation Department of Disease Pharmacology, LEO Pharma, Ballerup, Denmark

Hanne Norsgaard

Affiliation Department of Molecular Biomedicine, LEO Pharma, Ballerup, Denmark

Tord Labuda

Affiliation Department of Molecular Biomedicine, LEO Pharma, Ballerup, Denmark

Pieter Hupkens

Affiliation Department of Plastic Surgery Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Piet E. J. van Erp

Affiliation Dermatology Department, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Irma Joosten

Affiliation Laboratory of Medical Immunology, Department of Laboratory Medicine, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Hans J. P. M. Koenen

h.koenen@labgk.umcn.nl

Affiliation Laboratory of Medical Immunology, Department of Laboratory Medicine, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Competing Interests

Svensson L, Latta M, Norsgaard H, and Labuda T are all employees at LEO Pharma A/S. This does not alter our adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: VLO IJ HJPMK. Performed the experiments: VLO HN EF RRMCK. Analyzed the data: VLO HJPMK PEvE LS MMBS. Contributed reagents/materials/analysis tools: PH PCMvdK ML TL. Wrote the paper: VLO HJPMK IJ.