About the Authors

Yasusei Kudo

yasusei@tokushima-u.ac.jp (YK); ttakata@hiroshima-u.ac.jp (TT)

Affiliations Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan, Department of Oral Molecular Pathology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Shinji Iizuka

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Maki Yoshida

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Phuong Thao Nguyen

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Samadarani B. S. M. Siriwardena

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Takaaki Tsunematsu

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Mariko Ohbayashi

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Toshinori Ando

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Daijiro Hatakeyama

Affiliation Department of Oral and Maxillofacial Sciences, Gifu University Graduate School of Medicine, Gifu, Japan

Toshiyuki Shibata

Affiliation Department of Oral and Maxillofacial Sciences, Gifu University Graduate School of Medicine, Gifu, Japan

Keiichi Koizumi

Affiliation Department of Kampo Diagnostic, Institute of Natural Medicine, University of Toyama, Toyama, Japan

Masahiro Maeda

Affiliation Department of Research and Development, Immuno-Biological Laboratories, Co., Ltd., Fujioka-shi, Gunma, Japan

Naozumi Ishimaru

Affiliation Department of Oral Molecular Pathology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Ikuko Ogawa

Affiliation Center of Oral Clinical Examination, Hiroshima University Hospital, Hiroshima, Japan

Takashi Takata

yasusei@tokushima-u.ac.jp (YK); ttakata@hiroshima-u.ac.jp (TT)

Affiliation Department of Oral and Maxillofacial Pathobiology, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Competing Interests

Author Masahiro Maeda is employed by Immuno-Biological Laboratories, Co., Ltd. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YK. Performed the experiments: YK SI MY PTN SBSMS T. Tsunematsu MO TA. Analyzed the data: YK SI MY PTN NI IO T. Takata. Contributed reagents/materials/analysis tools: YK PTN SBSMS DH TS KK MM. Wrote the paper: YK. Obtained permission for use of cell line: KK.