About the Authors

John A. Hart

Affiliations Lukuru Wildlife Research Foundation, Kinshasa, Gombe, Democratic Republic of Congo, Division of Vertebrate Zoology, Yale Peabody Museum of Natural History, New Haven, Connecticut, United States of America

Kate M. Detwiler

kdetwile@fau.edu

Affiliation Department of Anthropology, Florida Atlantic University, Boca Raton, Florida, United States of America

Christopher C. Gilbert

Affiliations Department of Anthropology, Hunter College of the City University of New York, New York, New York, United States of America, New York Consortium in Evolutionary Primatology, New York, New York, United States of America

Andrew S. Burrell

Affiliation Center for the Study of Human Origins, Department of Anthropology, New York University, New York, New York, United States of America

James L. Fuller

Affiliations New York Consortium in Evolutionary Primatology, New York, New York, United States of America, Department of Ecology, Evolution and Environmental Biology, Columbia University, New York, New York, United States of America

Maurice Emetshu

Affiliation Lukuru Wildlife Research Foundation, Kinshasa, Gombe, Democratic Republic of Congo

Terese B. Hart

Affiliations Lukuru Wildlife Research Foundation, Kinshasa, Gombe, Democratic Republic of Congo, Division of Vertebrate Zoology, Yale Peabody Museum of Natural History, New Haven, Connecticut, United States of America

Ashley Vosper

Affiliation Wildlife Conservation Society Zanaga Project, Wildlife Conservation Society Congo, Brazzaville, Republic of Congo

Eric J. Sargis

Affiliations Division of Vertebrate Zoology, Yale Peabody Museum of Natural History, New Haven, Connecticut, United States of America, Department of Anthropology, Yale University, New Haven, Connecticut, United States of America

Anthony J. Tosi

Affiliation Center for the Study of Human Origins, Department of Anthropology, New York University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAH TBH KMD CCG ASB JLF AV EJS AJT. Contributed reagents/materials/analysis tools: JAH TBH KMD CCG ASB JLF ME AV EJS AJT. Wrote the paper: JAH KMD CCG ASB JLF TBH EJS AJT. Initial discovery and data collection: JAH TBH ME AV. Collected and analyzed the molecular data: JAH ME AV KMD ASB AJT. Collected and analyzed the morphological data: JAH ME AV CCG EJS. Collected and analyzed the behavioral and ecological data: JAH TBH AV JLF. Establishment of field site and proposed national park: JAH TBH ME.