About the Authors

Mizuho Ono

Affiliation Department of Medical Genetics, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Yuko Hamada

Affiliation Department of Medical Genetics, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Yasue Horiuchi

Affiliation Department of Psychiatry and Behavioral Sciences, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Mami Matsuo-Takasaki

Affiliation Department of Regenerative Medicine and Stem Cell Biology, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Yoshimasa Imoto

Affiliation Department of Otorhinolaryngology Head and Neck Surgery, Faculty of Medical Sciences, University of Fukui, Fukui City, Fukui, Japan

Kaishi Satomi

Affiliation Department of Diagnostic Pathology, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Tadao Arinami

Affiliation Department of Medical Genetics, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan

Mamoru Hasegawa

Affiliation DNAVEC Research Inc, Tsukuba, Ibaraki, Japan

Tsuyoshi Fujioka

Affiliation Cell Engineering Division, RIKEN BioResource Center, Tsukuba, Ibaraki, Japan

Yukio Nakamura

Affiliation Cell Engineering Division, RIKEN BioResource Center, Tsukuba, Ibaraki, Japan

Emiko Noguchi

enoguchi@md.tsukuba.ac.jp

Affiliations Department of Medical Genetics, Faculty of Medicine, University of Tsukuba, Tsukuba, Ibaraki, Japan, Japan Science and Technology Agency, Core Research for Evolutional Science and Technology (CREST), Chiyoda, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Y. Horiuchi MMT TA EN. Performed the experiments: MO Y. Hamada YI KS TF. Analyzed the data: MO TH EN. Contributed reagents/materials/analysis tools: MMT YN MH. Wrote the paper: MO Y. Hamada MMT EN.