About the Authors

Jorg J. M. Massen

jorgmassen@gmail.com

Affiliations Department of Behavioural Biology, Utrecht University, Utrecht, The Netherlands, Department of Cognitive Biology, University of Vienna, Vienna, Austria

Dorith A. Vermunt

Affiliation Department of Behavioural Biology, Utrecht University, Utrecht, The Netherlands

Elisabeth H. M. Sterck

Affiliations Department of Behavioural Biology, Utrecht University, Utrecht, The Netherlands, Ethology Research, Biomedical Primate Research Centre, Rijswijk, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JM. Performed the experiments: JM DV. Analyzed the data: JM. Contributed reagents/materials/analysis tools: JM. Wrote the paper: JM DV ES.