About the Authors

Madelyn M. Gerber

Affiliation Integrated Biomedical Sciences Graduate Program, The Ohio State University, Columbus, Ohio, United States of America

Heather Hampel

Affiliations The Ohio State University College of Medicine, Columbus, Ohio, United States of America, Division of Human Genetics, Department of Internal Medicine, The Ohio State University, Columbus, Ohio, United States of America, Human Cancer Genetics Program, Comprehensive Cancer Center, The Ohio State University, Columbus, Ohio, United States of America

Nathan P. Schulz

Affiliation The Ohio State University College of Medicine, Columbus, Ohio, United States of America

Soledad Fernandez

Affiliation The Center for Biostatistics, The Ohio State University, Columbus, Ohio, United States of America

Lai Wei

Affiliation The Center for Biostatistics, The Ohio State University, Columbus, Ohio, United States of America

Xiao-Ping Zhou

Affiliation Department of Pathology, The Ohio State University, Columbus, Ohio, United States of America

Albert de la Chapelle

Affiliations Division of Human Genetics, Department of Internal Medicine, The Ohio State University, Columbus, Ohio, United States of America, Human Cancer Genetics Program, Comprehensive Cancer Center, The Ohio State University, Columbus, Ohio, United States of America, Department of Molecular Virology, Immunology and Medical Genetics, The Ohio State University, Columbus, Ohio, United States of America

Amanda Ewart Toland

Amanda.Toland@osumc.edu

Affiliations Division of Human Genetics, Department of Internal Medicine, The Ohio State University, Columbus, Ohio, United States of America, Human Cancer Genetics Program, Comprehensive Cancer Center, The Ohio State University, Columbus, Ohio, United States of America, Department of Molecular Virology, Immunology and Medical Genetics, The Ohio State University, Columbus, Ohio, United States of America

Competing Interests

Amanda Toland is a PLoS ONE Editorial Board member. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MMG AET. Performed the experiments: MMG NPS. Analyzed the data: MMG NPS SF LW AET. Contributed reagents/materials/analysis tools: HH XPZ SF LW AdlC NPS AET. Wrote the paper: MMG AET. Edited manuscript: HH XPZ LW SF AdlC NPS.