About the Authors

Else Verbeek

else.verbeek@csiro.au

Affiliations Department of Biological Sciences, University of Waikato, Hamilton, New Zealand, Animal Behaviour and Welfare, AgResearch Limited, Hamilton, New Zealand

Mark Hope Oliver

Affiliation The Liggins Institute, University of Auckland, Auckland, New Zealand

Joseph Rupert Waas

Affiliation Department of Biological Sciences, University of Waikato, Hamilton, New Zealand

Lance Maxwell McLeay

Affiliation Department of Biological Sciences, University of Waikato, Hamilton, New Zealand

Dominique Blache

Affiliation School of Animal Biology, University of Western Australia, Crawley, Western Australia, Australia

Lindsay Ross Matthews

Affiliation Animal Behaviour and Welfare, AgResearch Limited, Hamilton, New Zealand

Competing Interests

The study received combined funding from Beef+Lamb NZ and Meat and Livestock Australia, as well as additional funding from the Foundation for Research, Science and Technology. Furthermore, two authors were employed by AgResearch, a New Zealand crown research institute. The funding sources and author employment by AgResearch do not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: EV MHO LRM JRW LMM. Performed the experiments: EV. Analyzed the data: EV. Contributed reagents/materials/analysis tools: DB MHO LRM. Wrote the paper: EV JRW MHO LRM DB LMM.