About the Authors

Piyachat Chansela

Affiliations Faculty of Science, Department of Anatomy, Mahidol University, Bangkok, Thailand, Department of Cell Biology and Anatomy, School of Medicine, Hamamatsu University, Hamamatsu, Shizuoka, Japan

Naoko Goto-Inoue

Affiliation Graduate School of Health Promotion Sciences, Tokyo Metropolitan University, Hachioji, Tokyo, Japan

Nobuhiro Zaima

Affiliation Department of Applied Biological Chemistry, Kinki University, Nara, Japan

Takahiro Hayasaka

Affiliation Department of Cell Biology and Anatomy, School of Medicine, Hamamatsu University, Hamamatsu, Shizuoka, Japan

Morakot Sroyraya

Affiliations Faculty of Science, Department of Anatomy, Mahidol University, Bangkok, Thailand, Department of Cell Biology and Anatomy, School of Medicine, Hamamatsu University, Hamamatsu, Shizuoka, Japan

Napamanee Kornthong

Affiliation Faculty of Science, Department of Anatomy, Mahidol University, Bangkok, Thailand

Attakorn Engsusophon

Affiliation Faculty of Science, Department of Anatomy, Mahidol University, Bangkok, Thailand

Montakan Tamtin

Affiliation Department of Fisheries, Coastal Aquatic Feed Research Institute, Coastal Fisheries Research and Development Bureau, Petchaburi, Thailand

Chatchawalee Chaisri

Affiliation Department of Fisheries, Coastal Aquatic Feed Research Institute, Coastal Fisheries Research and Development Bureau, Petchaburi, Thailand

Prasert Sobhon

Affiliation Faculty of Science, Department of Anatomy, Mahidol University, Bangkok, Thailand

Mitsutoshi Setou

setou@hama-med.ac.jp

Affiliation Department of Cell Biology and Anatomy, School of Medicine, Hamamatsu University, Hamamatsu, Shizuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PC N-GI NZ TH. Performed the experiments: PC. Analyzed the data: PC. Contributed reagents/materials/analysis tools: PS M. Setou. Wrote the paper: PC. Helped design the experiments and acted as technical consultants: NZ TH. Assisted in collecting the specimens, carried out the histological experiment, and provided technical support: M. Sroyraya AE NK. Cultured the shrimp: MT CC. Edited the manuscript and supervised the project: PS M. Setou.