About the Authors

Atsushi Ugajin

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Taketoshi Kiya

Affiliation Division of Life Sciences, Graduate School of Natural Science and Technology, Kanazawa University, Kanazawa, Japan

Takekazu Kunieda

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Masato Ono

stkubo@biol.s.u-tokyo.ac.jp (TK); ono@agr.tamagawa.ac.jp (MO)

Affiliation Laboratory of Applied Entomology and Zoology, Graduate School of Agriculture, Tamagawa University, Machida, Japan

Tadaharu Yoshida

Affiliation Honeybee Science Research Center, Tamagawa University, Machida, Japan

Takeo Kubo

stkubo@biol.s.u-tokyo.ac.jp (TK); ono@agr.tamagawa.ac.jp (MO)

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AU T. Kiya. Performed the experiments: AU T. Kiya MO TY. Analyzed the data: AU T. Kiya. Contributed reagents/materials/analysis tools: AU T. Kiya MO TY. Wrote the paper: AU T. Kunieda T. Kubo.