About the Authors

Shannon R. Matzinger

Affiliations Center for Comparative Medicine, University of California Davis, Davis, California, United States of America, California National Primate Research Center, University of California Davis, Davis, California, United States of America

Timothy D. Carroll

Affiliations Center for Comparative Medicine, University of California Davis, Davis, California, United States of America, California National Primate Research Center, University of California Davis, Davis, California, United States of America

Linda Fritts

Affiliations Center for Comparative Medicine, University of California Davis, Davis, California, United States of America, California National Primate Research Center, University of California Davis, Davis, California, United States of America

Michael B. McChesney

Affiliation Center for Comparative Medicine, University of California Davis, Davis, California, United States of America

Christopher J. Miller

cjmiller@ucdavis.edu

Affiliations Center for Comparative Medicine, University of California Davis, Davis, California, United States of America, California National Primate Research Center, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CJM MBM. Performed the experiments: SRM TDC LF. Analyzed the data: CJM SRM. Wrote the paper: CJM SRM.