About the Authors

Denver W. Fowler

df9465@yahoo.co.uk

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Elizabeth A. Freedman

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

John B. Scannella

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Robert E. Kambic

Affiliation Department of Ecology and Evolutionary Biology, Brown University, Providence, Rhode Island, United States of Ameria

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DWF. Performed the experiments: DWF EAF JBS. Analyzed the data: DWF EAF. Contributed reagents/materials/analysis tools: DWF EAF JBS REK. Wrote the paper: DWF EAF.