About the Authors

Mizue Hisano

Mizue.Hisano@jcu.edu.au

Affiliation School of Marine and Tropical Biology, James Cook University, Townsville, Australia

Sean R. Connolly

Affiliations School of Marine and Tropical Biology, James Cook University, Townsville, Australia, ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, Australia

William D. Robbins

Affiliation School of Marine and Tropical Biology, James Cook University, Townsville, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: MH. Wrote the paper: MH SRC. Obtained the data used in analysis: WDR. Conceived and designed the modeling method: MH SRC.