About the Authors

Abdulaziz M. Al-Swailem

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Maher M. Shehata

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Faisel M. Abu-Duhier

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Essam J. Al-Yamani

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Khalid A. Al-Busadah

Affiliation Faculty of Veterinary Medicine and Animal Resources, King Faisal University, Al-Hassa, Saudi Arabia

Mohammed S. Al-Arawi

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Ali Y. Al-Khider

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Abdullah N. Al-Muhaimeed

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Fahad H. Al-Qahtani

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Manee M. Manee

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Badr M. Al-Shomrani

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Saad M. Al-Qhtani

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Amer S. Al-Harthi

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Kadir C. Akdemir

Affiliation Department of Medicine, BIDMC Genomics Center, Harvard Medical School, Boston, Massachusetts, United States of America

Mehmet S. Inan

† Deceased.

Affiliation Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia

Hasan H. Otu

hotu@bidmc.harvard.edu

Affiliations Biotechnology Research Center, Natural Resources and Environment Research Institute, King Abdulaziz City for Science and Technology, Riyadh, Saudi Arabia, Department of Medicine, BIDMC Genomics Center, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AMAS MMS FMAD EJAY KAAB MSAA AYAK ANAM FHAQ MMM BMAS SMAQ ASAH MSI HHO. Performed the experiments: AMAS MMS EJAY MSAA AYAK MSI. Analyzed the data: MSAA ANAM FHAQ MMM BMAS SMAQ ASAH KCA HHO. Contributed reagents/materials/analysis tools: AMAS MMS EJAY ANAM FHAQ MMM BMAS SMAQ ASAH KCA MSI. Wrote the paper: AMAS MMS FMAD EJAY KAAB AYAK KCA MSI HHO.