About the Authors

Ken Tsumiyama

Affiliation Department of Biophysics, Kobe University Graduate School of Health Science, Kobe, Japan

Yumi Miyazaki

Affiliation Department of Biophysics, Kobe University Graduate School of Health Science, Kobe, Japan

Shunichi Shiozawa

shioz@med.kobe-u.ac.jp

Affiliations Department of Biophysics, Kobe University Graduate School of Health Science, Kobe, Japan, Department of Medicine, Kobe University Graduate School of Medicine, Kobe, Japan, The Center for Rheumatic Diseases, Kobe University Hospital, Kobe, Japan, Global Center of Excellence (GCOE), Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS. Performed the experiments: KT YM. Analyzed the data: KT YM SS. Wrote the paper: KT SS.