About the Authors

Hiroshi Tomita

hiroshi-tomita@iiare.tohoku.ac.jp

Affiliation International Advanced Interdisciplinary Research, Tohoku University, Sendai, Japan

Eriko Sugano

Affiliation International Advanced Interdisciplinary Research, Tohoku University, Sendai, Japan

Yugo Fukazawa

Affiliations Division of Cerebral Structure, National Institute for Physiological Sciences, Okazaki, Japan, Sokendai, Kanagawa, Japan

Hitomi Isago

Affiliations International Advanced Interdisciplinary Research, Tohoku University, Sendai, Japan, Department of Medical Biochemistry, Tohoku University Graduate School of Medicine, Sendai, Japan

Yuka Sugiyama

Affiliations Tohoku University Basic and Translational Research Center for Global Brain Science, Sendai, Japan, Department of Physiology and Pharmacology, Tohoku University Graduate School of Medicine, Sendai, Japan

Teru Hiroi

Affiliation International Advanced Interdisciplinary Research, Tohoku University, Sendai, Japan

Toru Ishizuka

Affiliation Department of Developmental Biology and Neuroscience, Tohoku University Graduate School of Life Sciences, Sendai, Japan

Hajime Mushiake

Affiliation Department of Physiology, Tohoku University Graduate School of Medicine, Sendai, Japan

Megumi Kato

Affiliation Section of Mammalian Transgenesis, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Japan

Masumi Hirabayashi

Affiliations Sokendai, Kanagawa, Japan, Section of Mammalian Transgenesis, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Japan

Ryuichi Shigemoto

Affiliations Division of Cerebral Structure, National Institute for Physiological Sciences, Okazaki, Japan, Sokendai, Kanagawa, Japan, Solution-Oriented Research for Science and Technology, Kawaguchi, Japan

Hiromu Yawo

Affiliations Department of Developmental Biology and Neuroscience, Tohoku University Graduate School of Life Sciences, Sendai, Japan, Tohoku University Basic and Translational Research Center for Global Brain Science, Sendai, Japan

Makoto Tamai

Affiliation Department of Medical Biochemistry, Tohoku University Graduate School of Medicine, Sendai, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HT ES YF TI RS. Performed the experiments: HT ES YF HI YS TH MK MH. Analyzed the data: HT. Contributed reagents/materials/analysis tools: HM. Wrote the paper: HT HY MT.