About the Authors

Elena S. Suvorova

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

Olivier Lucas

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

Carla M. Weisend

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

MaryClare F. Rollins

Affiliation Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America

Gary F. Merrill

Affiliation Biochemistry and Biophysics, Oregon State University, Corvallis, Oregon, United States of America

Mario R. Capecchi

Affiliation Howard Hughes Medical Institute (HHMI), University of Utah, Salt Lake City, Utah, United States of America

Edward E. Schmidt

eschmidt@montana.edu

Affiliations Veterinary Molecular Biology, Montana State University, Bozeman, Montana, United States of America, Center for Reproductive Biology, Washington State University, Pullman, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ESS EES. Performed the experiments: ESS CMW MFR EES. Analyzed the data: ESS OL EES. Contributed reagents/materials/analysis tools: GM MRC EES. Wrote the paper: ESS EES.