About the Authors

Jie Pan

Contributed equally to this work with: Jie Pan, Jin Wang

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Jin Wang

Contributed equally to this work with: Jie Pan, Jin Wang

Affiliation Key Laboratory for Nuclear Waste Treatment and Environmental Safety, Commission of Science, Technology and Industry for National Defence, Southwest University of Science and Technology, Mianyang, China

Zhengfu Zhou

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Yongliang Yan

Affiliations Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China, National Centre for Plant Gene Research, Beijing, China

Wei Zhang

Affiliations Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China, National Centre for Plant Gene Research, Beijing, China

Wei Lu

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Shuzhen Ping

Affiliation National Centre for Plant Gene Research, Beijing, China

Qilin Dai

Affiliation Key Laboratory for Nuclear Waste Treatment and Environmental Safety, Commission of Science, Technology and Industry for National Defence, Southwest University of Science and Technology, Mianyang, China

Menglong Yuan

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Bin Feng

Affiliation Key Laboratory for Nuclear Waste Treatment and Environmental Safety, Commission of Science, Technology and Industry for National Defence, Southwest University of Science and Technology, Mianyang, China

Xiaoguang Hou

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Ying Zhang

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Ma Ruiqiang

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Tingting Liu

Affiliation Key Laboratory for Nuclear Waste Treatment and Environmental Safety, Commission of Science, Technology and Industry for National Defence, Southwest University of Science and Technology, Mianyang, China

Lu Feng

Affiliation TEDA School of Biological Sciences and Biotechnology and Tianjin Key Laboratory of Microbial Functional Genomics, Nankai University, Tianjin, China

Lei Wang

Affiliation TEDA School of Biological Sciences and Biotechnology and Tianjin Key Laboratory of Microbial Functional Genomics, Nankai University, Tianjin, China

Ming Chen

chenmingbio@hotmail.com (MC); linmin57@vip.163.com (ML)

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China

Min Lin

chenmingbio@hotmail.com (MC); linmin57@vip.163.com (ML)

Affiliations Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Key Laboratory of Crop Biotechnology, Ministry of Agriculture, Beijing, China, National Centre for Plant Gene Research, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LW MC ML. Performed the experiments: JP JW ZZ WZ SP QD BF XH YZ RM TL LF. Analyzed the data: JP JW YY WL MY MC ML. Wrote the paper: JW LW MC ML.