About the Authors

John G. Kenny

Affiliation School of Biological Sciences, University of Liverpool, Liverpool, United Kingdom

Deborah Ward

Affiliation School of Biological Sciences, University of Liverpool, Liverpool, United Kingdom

Elisabet Josefsson

Affiliation Department of Rheumatology and Inflammation Research, University of Gothenburg, Göteborg, Sweden

Ing-Marie Jonsson

Affiliation Department of Rheumatology and Inflammation Research, University of Gothenburg, Göteborg, Sweden

Jason Hinds

Affiliation Division of Cellular & Molecular Medicine, St George's, University of London, London, United Kingdom

Huw H. Rees

Affiliation School of Biological Sciences, University of Liverpool, Liverpool, United Kingdom

Jodi A. Lindsay

Affiliation Division of Cellular & Molecular Medicine, St George's, University of London, London, United Kingdom

Andrej Tarkowski

Affiliation Department of Rheumatology and Inflammation Research, University of Gothenburg, Göteborg, Sweden

Malcolm J. Horsburgh

M.J.Horsburgh@Liverpool.ac.uk

Affiliation School of Biological Sciences, University of Liverpool, Liverpool, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JGK DW EJ AT MJH. Performed the experiments: JGK DW EJ IMJ MJH. Analyzed the data: JGK DW EJ IMJ JH MJH. Contributed reagents/materials/analysis tools: JL. Wrote the paper: JGK DW HR MJH.