About the Authors

Hirotaka Sakamoto

hsakamo@koto.kpu-m.ac.jp

Affiliation Department of Anatomy and Neurobiology, Kyoto Prefectural University of Medicine, Kawaramachi-Hirokoji, Kamigyo-ku, Kyoto, Japan

Ken-Ichi Matsuda

Affiliation Department of Anatomy and Neurobiology, Kyoto Prefectural University of Medicine, Kawaramachi-Hirokoji, Kamigyo-ku, Kyoto, Japan

Damian G. Zuloaga

Affiliation Program in Neuroscience, Departments of Psychology and Zoology, Michigan State University, East Lansing, Michigan, United States of America

Nobuko Nishiura

Affiliation Department of Anatomy and Neurobiology, Kyoto Prefectural University of Medicine, Kawaramachi-Hirokoji, Kamigyo-ku, Kyoto, Japan

Keiko Takanami

Affiliation Department of Anatomy and Neurobiology, Kyoto Prefectural University of Medicine, Kawaramachi-Hirokoji, Kamigyo-ku, Kyoto, Japan

Cynthia L. Jordan

Affiliation Program in Neuroscience, Departments of Psychology and Zoology, Michigan State University, East Lansing, Michigan, United States of America

S. Marc Breedlove

Affiliation Program in Neuroscience, Departments of Psychology and Zoology, Michigan State University, East Lansing, Michigan, United States of America

Mitsuhiro Kawata

Affiliation Department of Anatomy and Neurobiology, Kyoto Prefectural University of Medicine, Kawaramachi-Hirokoji, Kamigyo-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS MK. Performed the experiments: HS KIM DGZ NN KT. Analyzed the data: HS. Contributed reagents/materials/analysis tools: CLJ MB. Wrote the paper: HS MB MK.