About the Authors

Yufeng Shen

Affiliations Department of Computer Science, Columbia University, New York, New York, United States of America, Center for Computational Biology and Bioinformatics, Columbia University, New York, New York, United States of America

Sumeet Sarin

Affiliation Department of Biochemistry and Molecular Biophysics, Howard Hughes Medical Institute, Columbia University Medical Center, New York, New York, United States of America

Ye Liu

Affiliation Department of Computer Science, Columbia University, New York, New York, United States of America

Oliver Hobert

or38@columbia.edu (OH); ip2169@columbia.edu (IP)

Affiliation Department of Biochemistry and Molecular Biophysics, Howard Hughes Medical Institute, Columbia University Medical Center, New York, New York, United States of America

Itsik Pe'er

or38@columbia.edu (OH); ip2169@columbia.edu (IP)

Affiliations Department of Computer Science, Columbia University, New York, New York, United States of America, Center for Computational Biology and Bioinformatics, Columbia University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS SS YL OH IP. Performed the experiments: YS SS YL IP. Analyzed the data: YS SS YL OH IP. Wrote the paper: YS SS OH IP.