About the Authors

Robin M. D. Beck

robin.beck@student.unsw.edu.au

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Henk Godthelp

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Vera Weisbecker

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Michael Archer

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Suzanne J. Hand

Affiliation School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: RB HG VW. Wrote the paper: RB HG MA SH. Other: Collected and identified specimens: HG. Undertook initial analysis: HG. Interpretation of specimens: HG. Provided geological data: HG. Wrote parts of the supplementary information: HG. Carried out CT scanning, and processing and interpretation of CT images: VW. Oversaw project: MA. Supervised RMDB: MA SH. Provided biocorrelative data for supplementary information: SH. Helped write supplementary information: SH. Carried out all anatomical comparisons: RB. Carried out all analyses: RB. Wrote the majority of the paper and supplementary material: RB.