About the Authors

Michiel van Boven

*E-mail: michiel.van.boven@rivm.nl

Affiliations Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands, Animal Sciences Group, Wageningen University and Research Centre, Lelystad, The Netherlands, Centre for Ecological and Evolutionary Studies, University of Groningen, Haren, The Netherlands

Don Klinkenberg

Affiliation Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Ido Pen

Affiliation Centre for Ecological and Evolutionary Studies, University of Groningen, Haren, The Netherlands

Franz J. Weissing

Affiliation Centre for Ecological and Evolutionary Studies, University of Groningen, Haren, The Netherlands

Hans Heesterbeek

Affiliation Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HH FW DK Mv IP. Performed the experiments: Mv. Analyzed the data: Mv. Wrote the paper: HH FW DK Mv IP.