About the Authors

Drew T. Cronin

dtc33@drexel.edu

Affiliations Department of Biology, Drexel University, Philadelphia, Pennsylvania, United States of America, Department of Biodiversity, Earth and Environmental Science, Drexel University, Philadelphia, Pennsylvania, United States of America, Bioko Biodiversity Protection Program, Malabo, Bioko Norte, Equatorial Guinea

Stephen Woloszynek

Affiliations Department of Biology, Drexel University, Philadelphia, Pennsylvania, United States of America, Department of Electrical and Computer Engineering, Drexel University, Philadelphia, Pennsylvania, United States of America

Wayne A. Morra

Affiliation School of Global Business, Arcadia University, Glenside, Pennsylvania, United States of America

Shaya Honarvar

Affiliations Department of Biology, Drexel University, Philadelphia, Pennsylvania, United States of America, Department of Biology, Indiana University - Purdue University Fort Wayne, Fort Wayne, Indiana, United States of America

Joshua M. Linder

Affiliation Department of Sociology and Anthropology, James Madison University, Harrisonburg, Virginia, United States of America

Mary Katherine Gonder

Affiliations Department of Biology, Drexel University, Philadelphia, Pennsylvania, United States of America, Bioko Biodiversity Protection Program, Malabo, Bioko Norte, Equatorial Guinea

Michael P. O’Connor

Affiliations Department of Biology, Drexel University, Philadelphia, Pennsylvania, United States of America, Department of Biodiversity, Earth and Environmental Science, Drexel University, Philadelphia, Pennsylvania, United States of America, Bioko Biodiversity Protection Program, Malabo, Bioko Norte, Equatorial Guinea

Gail W. Hearn

Affiliations Department of Biology, Drexel University, Philadelphia, Pennsylvania, United States of America, Department of Biodiversity, Earth and Environmental Science, Drexel University, Philadelphia, Pennsylvania, United States of America, Bioko Biodiversity Protection Program, Malabo, Bioko Norte, Equatorial Guinea

Competing Interests

The authors have read the journal's policy and declare the following competing interests: Mobil Equatorial Guinea, Inc (MEGI). However, MEGI did not have any additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. The support received from MEGI does not alter the authors adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DTC WAM GWH. Performed the experiments: DTC WAM GWH. Analyzed the data: DTC SW WAM SH JML MKG MPO GWH. Contributed reagents/materials/analysis tools: DTC SW WAM MPO. Wrote the paper: DTC SW WAM SH JML MKG MPO GWH.