About the Authors

Stephen J. Price

s.price@ucl.ac.uk (SJP); a.griffiths3@exeter.ac.uk (AGFG)

Affiliations UCL Genetics Institute, University College London, Darwin Building, Gower Street, London, United Kingdom, Institute of Zoology, Zoological Society of London, London, United Kingdom

Trenton W. J. Garner

Affiliation Institute of Zoology, Zoological Society of London, London, United Kingdom

Francois Balloux

Affiliation UCL Genetics Institute, University College London, Darwin Building, Gower Street, London, United Kingdom

Chris Ruis

Affiliation UCL Genetics Institute, University College London, Darwin Building, Gower Street, London, United Kingdom

Konrad H. Paszkiewicz

Affiliation Wellcome Trust Biomedical Informatics Hub, Biosciences, Geoffrey Pope Building, University of Exeter, Streatham Campus, Exeter, United Kingdom

Karen Moore

Affiliation Wellcome Trust Biomedical Informatics Hub, Biosciences, Geoffrey Pope Building, University of Exeter, Streatham Campus, Exeter, United Kingdom

Amber G. F. Griffiths

s.price@ucl.ac.uk (SJP); a.griffiths3@exeter.ac.uk (AGFG)

Current address: FoAM Kernow, Studio E, Jubilee Warehouse, Commercial Road, Penryn TR10 8FG, United Kingdom (Current address)

Affiliation Environment and Sustainability Institute, University of Exeter, Penryn Campus, Penryn, Cornwall, United Kingdom

Competing Interests

AGFG is a member of the PLOS ONE editorial board. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AGFG TWJG FB. Performed the experiments: CR AGFG SJP TWJG. Analyzed the data: KHP KM CR SJP FB. Contributed reagents/materials/analysis tools: KHP KM. Wrote the paper: SJP AGFG.