About the Authors

Helen Titmarsh

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Scott Kilpatrick

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Jennifer Sinclair

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Alisdair Boag

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Elizabeth F. Bode

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Stephanie M. Lalor

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Donna Gaylor

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Jacqueline Berry

Affiliation Specialist Assay Laboratory (Vitamin D), Clinical Biochemistry, Manchester Royal Infirmary, Manchester, United Kingdom

Nicholas X. Bommer

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Danielle Gunn-Moore

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Nikki Reed

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Ian Handel

‡ These authors are joint senior authors on this work.

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Richard J. Mellanby

Richard.mellanby@ed.ac.uk

‡ These authors are joint senior authors on this work.

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RM IH. Performed the experiments: SK DGM NR NB JS SK EB AB DG HT RM IH JB SL. Analyzed the data: IH HT RM. Contributed reagents/materials/analysis tools: HT RM IH JB. Wrote the paper: SK DGM NR NB JS SK EB AB DG HT RM IH JB SL.