About the Authors

J. Jeffrey Root

Jeff.root@aphis.usda.gov

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Kevin T. Bentler

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Susan A. Shriner

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Nicole L. Mooers

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Kaci K. VanDalen

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Heather J. Sullivan

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Alan B. Franklin

Affiliation United States Department of Agriculture, National Wildlife Research Center, Fort Collins, CO, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJR SAS KKV ABF. Performed the experiments: JJR KTB NLM. Analyzed the data: JJR KTB NLM KKV HJS. Wrote the paper: JJR SAS KKV ABF.