About the Authors

William B. Monahan

Bill_Monahan@nps.gov

Affiliation National Park Service, Natural Resource Stewardship & Science, Fort Collins, Colorado, United States of America

Nicholas A. Fisichelli

Affiliation National Park Service, Natural Resource Stewardship & Science, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WBM NAF. Performed the experiments: WBM NAF. Analyzed the data: WBM NAF. Contributed reagents/materials/analysis tools: WBM NAF. Wrote the paper: WBM NAF.