About the Authors

Marie-Soleil Dubois

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Samantha Morello

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Kelsey Rayment

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Mark D. Markel

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Ray Vanderby Jr

Affiliation Department of Orthopedics & Rehabilitation, School of Medicine & Public Health, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Vicki L. Kalscheur

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Zhengling Hao

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Ronald P. McCabe

Affiliation Department of Orthopedics & Rehabilitation, School of Medicine & Public Health, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Patricia Marquis

Affiliations Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Gulfstream Park, Hallandale Beach, Florida, United States of America

Peter Muir

muirp@vetmed.wisc.edu

Affiliation Comparative Orthopaedic Research Laboratory, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Competing Interests

Please note that although Patricia Marquis is affiliated with the Gulfstream Park company in Florida, this does not alter the authors' adherence to PLOS ONE policies on sharing of data and materials.

Author Contributions

Conceived and designed the experiments: P. Muir MDM RV RPM. Performed the experiments: MSD SM KR VK ZH RPM P. Muir. Analyzed the data: MSD P. Muir. Wrote the paper: MSD SM P. Muir. Contributed limb specimens for analysis: P. Marquis.