About the Authors

Jeff A. Tracey

jatracey@usgs.gov

Affiliation U.S. Geological Survey, Western Ecological Research Center, San Diego Field Station, San Diego, California, United States of America

James Sheppard

Affiliation San Diego Zoo Institute for Conservation Research, Escondido, California, United States of America

Jun Zhu

Affiliation Department of Statistics and Department of Entomology, University of Wisconsin – Madison, Madison, Wisconsin, United States of America

Fuwen Wei

Affiliation Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Science, Beijing, People's Republic of China

Ronald R. Swaisgood

Affiliation San Diego Zoo Institute for Conservation Research, Escondido, California, United States of America

Robert N. Fisher

Affiliation U.S. Geological Survey, Western Ecological Research Center, San Diego Field Station, San Diego, California, United States of America

Competing Interests

Sempra Energy funded this study. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all of the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JAT JS JZ. Performed the experiments: JAT JS JZ FW. Analyzed the data: JAT JS. Contributed reagents/materials/analysis tools: JAT JS. Wrote the paper: JAT JS JZ FW RRS RNF. Field data: JS FW.