About the Authors

Stephen Buckley

Affiliation BioArCh, University of York, York, United Kingdom

Donatella Usai

Affiliation Istituto Italiano per l'Africa e l'Oriente, Roma; Centro Studi Sudanese e Sub-Sahariani, Treviso, Italy

Tina Jakob

Affiliation Department of Archaeology, Durham University, Durham, United Kingdom

Anita Radini

Affiliations BioArCh, University of York, York, United Kingdom, University of Leicester Archaeological Services (ULAS), School of Archaeology and Ancient History, University of Leicester, Leicester, United Kingdom

Karen Hardy

khardy@icrea.cat

Affiliation ICREA (Catalan Institution for Research and Advanced Studies), Departament de Prehistòria, Universitat Autònoma de Barcelona, Bellaterra, Spain

Competing Interests

Co-author Karen Hardy is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: KH SB DU. Performed the experiments: KH SB. Analyzed the data: KH SB AR. Contributed reagents/materials/analysis tools: KH SB TJ DU. Contributed to the writing of the manuscript: KH SB DU TJ AR. Director of the excavations at the site: DU. Sorted, identified and selected all the samples: TJ.