About the Authors

Yi Ta Shao

Affiliations Department of Zoology, Stockholm University, Stockholm, Sweden, Sensory Physiology Laboratory, Marine Research Station, Academia Sinica, I-lain, Taiwan

Feng-Yu Wang

Affiliation Taiwan Ocean Research Institute, National Applied Research Laboratories, Kaohsiung, Taiwan

Wen-Chun Fu

Affiliation Sensory Physiology Laboratory, Marine Research Station, Academia Sinica, I-lain, Taiwan

Hong Young Yan

Contributed equally to this work with: Hong Young Yan, Bertil Borg

bertil.borg@zoologi.su.se (BB); hyyan@gate.sinica.edu.tw (HYY)

Affiliations Sensory Physiology Laboratory, Marine Research Station, Academia Sinica, I-lain, Taiwan, Hanse-Wissenschaftskolleg Institute for Advanced Study, Delmenhorst, Germany

Kazuhiko Anraku

Affiliation Faculty of Fisheries, Kagoshima University, Kagoshima, Japan

I-Shiung Chen

Affiliation Institute of Marine Biology, National Taiwan Ocean University, Keelung, Taiwan

Bertil Borg

Contributed equally to this work with: Hong Young Yan, Bertil Borg

bertil.borg@zoologi.su.se (BB); hyyan@gate.sinica.edu.tw (HYY)

Affiliation Department of Zoology, Stockholm University, Stockholm, Sweden

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YTS HYY KA BB. Performed the experiments: YTS FYW WCF BB. Analyzed the data: YTS WCF. Contributed reagents/materials/analysis tools: HYY KA ISC BB. Wrote the paper: YTS HYY BB.